We used reliable macrofossils of Araliaceae from Dendropanax eocenensis identified from leaf material (Dilcher & Dolph, 1970) and Paleopanax (= Metapanax) identified from fruit material, collected from middle Eocene deposits in North America (Manchester, 1994).
The traditional circumscriptions of Araliaceae and Apiaceae and their relationships as a "family pair" (sister families) are not supported.
This genus was also thought to have affinities to Araliaceae (especially K.
Suborder Apiineae (Pittosporaceae, Araliaceae, Myodocarpaceae, and Apiaceae) originated c.
Radiations within a period of 15 Ma led to the origin of the four lineages that are currently recognized as families: Pittosporaceae, Araliaceae, Myodocarpaceae, and Apiaceae (Fig.
The analyses show that Araliaceae had a median minimum stem age c.
Apart from Hydrocotyle-Trachymene and Harmsiopanax, Afro-Madagascan Schefflera is the earliest diverging clade among the rest of Araliaceae, dating back to the early Eocene with a stem age of c.
The remaining groups of Araliaceae have a stem age of c.
Araliaceae has an extensive fossil record that goes back to the Late Cretaceous but that is in need of revision; few of the araliaceous fossils can be considered reliable and revising them would be of extreme importance.
Oreopanax dakotensis, a new species of the Araliaceae from the Paleocene of North Dakota.
Aquifoliaceae Ilex antiqua Knobloch et No Mai 1986 Ilex gonnensis Mai 1970 Yes Ilex hercynica Mai 1970 Yes Ilex No Ilexpollenites No Table 6 Early Fossil Record of the Apiales Fossil taxon Organ Age Torricelliaceae Torricellia bonesii Fruit Early Miocene (Manchester) Manchester 1999 Torricellia bonesii Fruit Middle Eocene (Manchester) Manchester 1999 Torricellia bonesii Fruit Middle Eocene (Manchester) Manchester 1999 Torricellia bonesii Fruit Middle Eocene (Manchester) Manchester 1999 Araliaceae ("Apiaceae") Hydrocotyle sp Lancucka- Fruit Miocene Srodoniowa 1979 Araliaceae Aralia antiqua Knobloch Endocarp Maastrichtian et Mai 1986 Aralia cf.