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The rate of origination of chitinozoan species increases considerably within this biozone (Nestor 2009b): 9-10 species appear in the Kolka, Ohesaare, Ruhnu and Kaugatuma cores, fewer in the others.
The succession begins with the upper-most part of the lower Wenlock chitinozoan Interzone IV, established between the Margachitina margaritana and Conochitina manilla biozones (V.
Their opinion was based on lithological correlation with sections on the Estonian mainland and the stratigraphical distribution of the chitinozoan Cyathochitina regnelli sensu Nolvak & Grahn (1993).
For example, those of the ostracodes Plicibeyrichia numerosa-Undulirete baltica (Sarv 1982; Gailite 1986); the thelodont Thelodus sculptilis (Marss 1997); the conodont Ozarkodina crispa (Brazauskas 1993; Viira 1999); and the chitinozoan Eisenackitina barrandei (Nestor 2009).
The first attempt to divide the upper Silurian of the Ohesaare core section on the basis of chitinozoan assemblages was made by Nestor (1976).
Those of seven chitinozoan biozones are presented according to Nestor (2009, 2011, 2012), seven conodont ones from Viira & Aldridge (1998) and Viira (1999, 2000) and seven vertebrate indices according to Marss (1986, 1997) and Marss & Mannik (2013).
Precise time-correlation in the Ordovician can be achieved by biostratigraphical methods, particularly using graptolite, conodont and chitinozoan distribution (biozones).
In the chitinozoan biozonation, the Eisenackitina rhenana Subzone of the Laufeldochitina stentor Zone corresponds to the lower part of the Kukruse Stage (Nolvak & Grahn 1993; Nolvak 1997).