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ecosystem
Complex of living organisms, their physical environment, and all their interrelationships in a particular unit of space. An ecosystem's abiotic (nonbiological) constituents include minerals, climate, soil, water, sunlight, and all other nonliving elements; its biotic constituents consist of all its living members. Two major forces link these constituents: the flow of energy and the cycling of nutrients. The fundamental source of energy in almost all ecosystems is radiant energy from the sun; energy and organic matter are passed along an ecosystem's food chain. The study of ecosystems became increasingly sophisticated in the later 20th century; it is now instrumental in assessing and controlling the environmental effects of agricultural development and industrialization. See also biome.
ecosystem
ecosystem [′ek·o‚sis·təmor′ē·kō‚sis·təm]
Ecosystem
A functional system that includes an ecological community of organisms together with the physical environment, interacting as a unit. Ecosystems are characterized by flow of energy through food webs, production and degradation of organic matter, and transformation and cycling of nutrient elements. This production of organic molecules serves as the energy base for all biological activity within ecosystems. The consumption of plants by herbivores (organisms that consume living plants or algae) and detritivores (organisms that consume dead organic matter) serves to transfer energy stored in photosynthetically produced organic molecules to other organisms. Coupled to the production of organic matter and flow of energy is the cycling of elements. See Ecological communities, Environment
All biological activity within ecosystems is supported by the production of organic matter by autotrophs (organisms that can produce organic molecules such as glucose from inorganic carbon dioxide; see illustration). More than 99% of autotrophic production on Earth is through photosynthesis by plants, algae, and certain types of bacteria. Collectively these organisms are termed photoautotrophs (autotrophs that use energy from light to produce organic molecules). In addition to photosynthesis, some production is conducted by chemoautotrophic bacteria (autotrophs that use energy stored in the chemical bonds of inorganic molecules such as hydrogen sulfide to produce organic molecules). The organic molecules produced by autotrophs are used to support the organism's metabolism and reproduction, and to build new tissue. This new tissue is consumed by herbivores or detritivores, which in turn are ultimately consumed by predators or other detritivores.
Terrestrial ecosystems, which cover 30% of the Earth's surface, contribute a little over one-half of the total global photosynthetic production of organic matter—approximately 60 × 1015 grams of carbon per year. Oceans, which cover 70% of the Earth's surface, produce approximately 51 × 1015 g C y-1 of organic matter. See Biomass
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Food webs
Organisms are classified based upon the number of energy transfers through a food web (see illustration). Photoautotrophic production of organic matter represents the first energy transfer in ecosystems and is classified as primary production. Consumption of a plant by a herbivore is the second energy transfer, and thus herbivores occupy the second trophic level, also known as secondary production. Consumer organisms that are one, two, or three transfers from photoautotrophs are classified as primary, secondary, and tertiary consumers. Moving through a food web, energy is lost during each transfer as heat, as described by the second law of thermodynamics. Consequently, the total number of energy transfers rarely exceeds four or five; with energy loss during each transfer, little energy is available to support organisms at the highest levels of a food web. See Ecological energetics, Food web
Biogeochemical cycles
In contrast to energy, which is lost from ecosystems as heat, chemical elements (or nutrients) that compose molecules within organisms are not altered and may repeatedly cycle between organisms and their environment. Approximately 40 elements compose the bodies of organisms, with carbon, oxygen, hydrogen, nitrogen, and phosphorus being the most abundant. If one of these elements is in short supply in the environment, the growth of organisms can be limited, even if sufficient energy is available. In particular, nitrogen and phosphorus are the elements most commonly limiting organism growth. This limitation is illustrated by the widespread use of fertilizers, which are applied to agricultural fields to alleviate nutrient limitation. See Biogeochemistry, Nitrogen cycle
Carbon cycles between the atmosphere and terrestrial and oceanic ecosystems. This cycling results, in part, from primary production and decomposition of organic matter. Rates of primary production and decomposition, in turn, are regulated by the supply of nitrogen, phosphorus, and iron. The combustion of fossil fuels is a recent change in the global cycle that releases carbon that has long been buried within the Earth's crust to the atmosphere. Carbon dioxide in the atmosphere traps heat on the Earth's surface and is a major factor regulating the climate. This alteration of the global carbon cycle along with the resulting impact on the climate is a major issue under investigation by ecosystem ecologists. See Conservation of resources, Human ecology
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