As the tide is rising before high tide is the best time to see the hordes of black-tailed godwits
, as well as other waders such as knots, and ducks such as teals and pintails which also occur here in their thousands,'' he added.
A mated pair of black-tailed godwits
may fly off to separate wintering grounds a thousand kilometers apart, but they can return to their breeding grounds almost simultaneously, a migration survey has revealed.
Conservationists were concerned about the effect of dredging on ecology in the Dee estuary, one of the UK's most important areas for black-tailed godwits
and corn buntings were among the more unusual species which were spotted.
, a species of shorebirds, may be the fattest birds still able to get their lardy rumps into the air, but they could also be the avian marathon champs.
The research team was supported by a network of more than 2,000 birdwatchers who reported sightings of colour-ringed black-tailed godwits
as they travel from Iceland to Spain and Portugal.
Tunnicliffe's book Shorelands Summer Diary included 16 paintings of golden plovers, whimbrel, shoveler ducks, shag, black-tailed godwits
, red-necked phalarope and peregrine falcons.
Conservation authorities were concerned about the effect of dredging on ecology in the Dee estuary, one of the UKs most important areas for black-tailed godwits
An avocet - the emblem of the RSPB - has also called in at Saltholme this spring and the grassland has attracted a flock of rare black-tailed godwits
Other works feature redshanks in the golden glow of sunset on Budle Bay, a buzzard over Debdon at Rothbury, roe deer in birch woods and among cotton grass, a goshawk at Cragside, a barn owl at Hepple, puffins on the Farne Islands and a wheeling flock of bar-tailed godwits
Several green sandpipers were there over the weekend, a hard bird to see in North Wales, along with more than 30 black-tailed godwits
returning from Iceland.
But, though we are now waving goodbye to Wales' summer breeders at the reserve, over the coming months we can get ready to greet thousands upon thousands of birds that will spend the winter here - from dunlin, ringed plover and black-tailed godwits