Previous investigations in the study area revealed a stable presence of five predator families in small pools along the Jaribuni River in the Kenyan coastal province which were mainly in two orders: Hemiptera (Gerridae, Hydrometridae, Veliidae
and Notonectidae) and Coleoptera (Dytiscidae) (Mbogo, personal communication).
A phylogeny for all families and subfamilies, and for all genera but those assigned to the two most diverse families, Veliidae and Gerridae, is compiled from the many studies by the late Dr.
This study confirmed Mesoveliidae as being the sister group to all other families of Gerromorpha and also the close relationship between the Gerridae and Veliidae, but suggested that the superfamilies Gerroidea (Hermatobatidae + Gerridae + Veliidae) and Hydrometroidea (Paraphrynoveliidae + Macroveliidae + Hydrometridae), the family Veliidae, and the gerrid subfamily Cylindrostethinae were paraphyletic entities, and that Gerrinae, Haloveliinae, Ocelloveliinae, Paraphrynoveliidae, and Veliinae lacked convincing morphological diagnostic characters.
On the other hand, the Gerromorpha contain six well-supported and/or well-diagnosed clades: Mesoveliidae, Hebridae, Hermatobatidae, Hydrometridae, Macroveliidae + Paraphrynoveliidae and Veliidae + Gerridae, five of which were already recognized by Andersen (1982).
2009) found that the doublebipod gait in Hermatobates Carpenter, 1892 differs fundamentally from the syncronous rowing in members of the Halobatinae and Haloveliinae, which further separates the family from the Veliidae + Gerridae clade.
Damgaard (2008b) found Veliinae, Rhagoveliinae, Perittopinae, and Ocelloveliinae as succeeding sister taxa to the Gerridae + Haloveliinae + Microveliinae clade, but with poor support and diagnostic character combinations, except for Gerridae + Veliidae less Ocelloveliinae, which foremost could be diagnosed by the lack of ocelli in the macropterous morphs, although this character is found convergently in Madeoveliinae and Hydrometrinae + Limnobatodinae.
C4, C6 Trepobates pictus (Herrich-Schaeffer): B1, B3, C1 Trepobates subnitidus Esaki: B2, C2, C3 Family Mesoveliidae Mesovelia mulsanti White: A12, C4 Family Nepidae Ranatrafusca Palisot: A2, A8, A13 Family Notonectidae Notonecta irrorata Uhler: A11, B3, B4, C10 Family Pleidae Neoplea striola (Fieber): A2, B2, C5 Family Veliidae
Water-striders (Hemiptera: Gerridae, Veliidae, etc.
Marine Hetertoptera (Insecta): Gerridae, Veliidae, Hermatobatidae, Saldidae and Omaniidae, with notes on ecology and insular zoogeography.
Marine Heteroptera of the Northern Territory, including the first fresh-water species of Halobates Eschscholtz (Gerridae, Veliidae, Hermatobatidae and Corixidae).