In angiosperms (for gymnosperms, see Behnke, 1974), P-type plastids first appear in eumagnoliids (Winterales, Piperales, Magnoliales, Laurales), where, with the exception of
Himantandraceae, Lactoridaceae, Piperaceae, and Saururacene, one or several different forms (Plc, Pics, Plcfs, Plfs) are found in at least some species (Behnke, 1988).