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Sympetalae

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The following article is from The Great Soviet Encyclopedia (1979). It might be outdated or ideologically biased.

Sympetalae

 

a subclass of angiospermous dicotyledonous plants that includes families whose representatives are characterized by united petals. The petals may be united partially (only at the base or up to the middle of the petals) or completely.

In most Sympetalae the concresced basal part of the petals forms a tubule, and the upper parts resemble lobes, segments, or denticles. Also characteristic are a gamosepalous calyx, a cyclical flower, and one ovular covering (and not two, as in most polypet-alous plants). The staminal filaments and the floral tubule con-cresce over a more or less considerable surface.

The classification of the Sympetalae as a separate group (opposed to the Choripetalae) was proposed in 1864 by the German botanist A. Braun. In 1892 the German botanist A. Engler proposed the term “Metachlamydeae” for the Sympetalae because he considered the Sympetalae to be more highly organized than the Choripetalae and the Archichlamydeae. Most modern phylogenetic systems do not distinguish the Sympetalae as a separate subclass of dicotyledons.

M. E. KIRPICHNIKOV

The Great Soviet Encyclopedia, 3rd Edition (1970-1979). © 2010 The Gale Group, Inc. All rights reserved.
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References in periodicals archive
This group has been recognized by botanists since the eighteenth century, receiving names such as the Monopetalae, Gamopetalae or Sympetalae (Wagenitz, 1992) all of which allude to the characteristic connate corolla.
Velarde's papers followed the arrangement of Engler's classification system, but only listed a total of 700 names for several families (except those in the subclass Sympetalae), and thus did not include the Ericaceae.
Distribution of the character states "early sympetaly" and "late sympetaly" within the "Sympetalae Tetracyclicae" and presumably allied groups.
However, as Dahlgren (1940) correctly observed, there is a critical difference between a postament, which is a resistant nucellar protuberance into the embryo sac (the lateral parts having often been eaten away), and a small resistant group of nucellar cells not projecting into the embryo sac, for which he proposed the term "podium." A podium is therefore an insignificant structure, present in many plants with an insignificant nucellus, such as in Euphorbiaceae and many tenuinucellate Sympetalae (Dahlgren, 1940; Bor & Kapil, 1975).
Excluded Families: The Rubiaceae (Schumann, 1891) was placed in the Rubiales under the Sympetalae. The Convolvulaceae (including the Cuscutaceae as the subfamily Cuscutoideae) (Peter, 1891) was placed under the series Tubiflorae in the subseries Convolvulineae.
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