Teeth with multicuspid cladodont crown; central cusp twice as high and three times more broad than the lateral and intermediate cusps; pair of high accessory cusplets placed outside the lateral cusps; two rows of numerous high and small accessory cusplets placed labially; intermediate cusps and cusplets differently directed and incurved; labial cristae on the central cusp extended and prominent at the base; sub-lenticular base very elongated mesio-distally; ridge-like apical button considerably elongated, placed close to the lingual rim; shelf-like wide labio-basal projection; dense network of vascular canals occupying the entire tooth base.
The teeth possess a multicuspid cladodont crown made up of the main central, intermediate and lateral cusps and numerous intermediate and accessory cusplets.
The teeth have a cladodont crown with large central, moderate lateral and short intermediate cusps.
The diplodont, cladodont, phoebodont and protacrodont crown types appear in the elasmobranch dentitions in the Givetian.
Recently teeth of the oldest protacrodontid and cladodont (possibly symmoriid) chondrichthyans were discovered in the Late Givetian of the Kuznetsk Basin, western Siberia, Russia (Ivanov & Rodina 2007).