Similar reservations about the monophyly of Oedipodinae* (G-M plus Duroniella) can be expressed, given the presence of acridines within clades J, K and M and of Melanotettix within L.
All, except Pnorisa, are acridines (sensu OSF2) and all are African.
The acridines at the base of the tree had fairly ancient ancestors preceding the Gomphocerinae* --Oedipodinae* split by 10-70 my.
That is, pre-acridines, having spread throughout the conjoined African -South American continent, became separated into two groups postfragmentation, one group giving rise to the gomphocerine acridine complex that spread throughout South America and the other evolving into the oedipodine-acridine complex in Africa.
2007), remains the same with successive links to the acridines, Caledia and Froggattina.
Others have examined the phylogenetic position of Acrida with respect to a limited number of acridines, gomphocerines and oedipodines.
This clade (PP = 99%) diversified about 90 mya from a common neotropical ancestor and is external to A-D, consisting of a mixture of acridine and gomphocerine genera.
One led to the African acridine, Anaeolopus, another to the North American Stethophyma gracile and at least one migration established Aiolopus strepens throughout the Old World and Asia-Australia.