Subgenital plate as long as broad, with bristles evenly; sternal apodeme narrow, about 0.2 times longer than sternum (sternal apodeme plus sterno) (Figure 1C); genital capsule (Figures 1D, E) with paramere rounded apically, narrower than parameral lamina, with dense bristles; volsellar lamina with few bristles spaced subapically; cuspis robust, with tooth in lateral and apical parts and a bristle at the apex; digitus narrow, 0.40-0.46 times the length of cuspis plus volsellar lamina, with tooth in the apex, apex rounded dorsally and angulated ventrally, its basal region slightly larger than apical region; aedeagus
(including aedeagal apodeme) 0.90-0.95 times the length of paramere plus parameral lamina (including the parameral apodeme); ergot about 0.27-0.37 times the length of aedeagus
: In ventral view the medial lobe shows two sclerotized plates far apart from each other and tapering apicad, covering almost all of it.
Male terminalia: Hypoproct slightly bilobed, wider than cercus, with pair of posterior setae; aedeagus
longer than hypoproct and parameres; parameres apically with 2 to 3 short setae; gonocoxite cylindrical; gonostylus attenuate, widest at base, tapering to narrow apex, setulose basally, carinate beyond; apical tooth present.
The complexity of the phallic anatomy within the Serpusiae, and especially of the aedeagus
, was previously examined by Jago (1970), in his study of the genus Auloserpusia.
External male genitalia morphology in both species is distinguishable by the overall structure of genital capsule, shape of clasp, and aedeagus
. In D.
1E) with small stem, blade broad, oval-shaped, inner margin distinctly wavy, apex without spine and sub-round, outer upper margin near apex depressed distinctly, inner triangular spine present; inflated aedeagus
El material organico adyacente a las estructuras de la genitalia masculina (capsula genital y aedeagus
) fue eliminado.
Then, the male held the female with the pairs of legs, and positioned the posterior part of its body next to the female pygidium and inserted the aedeagus
to begin copulation.
1g): Symmetrical, Uncus sclerotised, elongated downward curved, slender at apex, tegumen short, broad dorsally, saccus V-shaped, valva elongated, basal part narrow, outer margin sinuated, inner margin sinuated with small hair, distally somewhat rounded lobe covered with hair, aedeagus
broad, long, curved, anteriorly narrow, posteriorly broad.
asymmetrical through 90[degrees] inversion of distiphallus; with two parameres, and these carrying a rugose morphoventral projection (Fig.
The most reliable characters for discrimination of Lilioceris species are those of the internal sac of the aedeagus
. In Criocerinae, these characters were used previously for separation of Lilioceris species from Iran and Oulema species related to Oulema melanopus L.