Asperula


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Asperula

 

(woodruff), a genus of annual and perennial herbs and subshrubs of the family Rubiaceae. The stems are square and smooth; the leaves are whorled or, rarely, opposite. The small, usually tetramerous flowers are most commonly in a panicle or head. The fruit is dry and separates into two sections.

There are about 90 species (according to other data, as many as 200), distributed in Eurasia, primarily in the Mediterranean. The USSR has 70 species. A. odorata (or Galium odoratum) is found in broad-leaved and mixed forests and in ravines. Dyer’s woodruff tinctoria), which contains a dye substance in its roots, grows in the European USSR and in the southern part of Western Siberia in meadows, on steppe slopes, and in pine forests. Many species are nectar-bearers. Several species, including A. azurea (or A. orientalis) and Asperula nítida, are cultivated as ornamentals. Some species, for example, A. odorata, are frequently included in the genus Galium.

T. V. EGOROVA

References in periodicals archive ?
Las especies constantes son Bulbostylis asperula (r), Poaceae sp.
trichostachyum 1, Asperula supina 1, Bupleurum falcatum subsp.
stevenii 5, Thymus cherlerioides 5, Alopecurus vaginatus 3, Bromopsis taurica 3, Paronychia cephalotes 3, Scorzonera crispa 3, Sideritis taurica 3, Thalictrum minus 3, Teucrium chamaedrys 3, Asperula supina 2, Elymus reflexiaristatus subsp.
ponticum, Asperula tenella, Asphodeline taurica, Centaurea orientalis, Convolvulus cantabrica, Crupina vulgaris, Galium biebersteinii, Onobrychis arenaria subsp.
Companions: Potentilla cinerea 3, Festuca gracilior 2, Sedum amplexicaule 2, Minuartia hybrida 2, Molineriella laevis 2, Jasione montana 1, Fumana procumbens 1, Helianthemum apenninum 1, Helianthemum incanum 1, Teucrium expassum 1, Saxifraga dichotoma 1, Herniaria cinerea 1, Leontodon longirostris 1, Pilosella officinarum 1, Asperula cynanchica 1, Calluna vulgaris +, Halimium viscosum +, Lavandula pedunculata +, Cerastium gracile 1, Corynephorus canescens +, Eryngium campestre +, Tuberaria guttata +.
cossoniana 3 y Galium fruticescens 4 en 4; Hieracium amplexicaule, Asperula cynanchica subsp.
Character taxa of the order (and of subordinate syntaxa): Achillea virescens, Argyrolobium zanonii, Allium sphaerocephalon, Arabis collina, Asperula purpurea, Avenula praetutiana, Bromus hordeaceus subsp.
Differential taxa of the alliance (against Filipendulo vulgaris-Helictotrichion pratensis): Anacamptis pyramidalis, Asperula cynanchica, Ctenidium molluscum, Festuca guestfalica subsp.
III Transgressive species from Festuco-Brometea: Cerastium arvense IV IV Asperula cynanchica II II Carlina acaulis subsp.