(2004), based on the same fossils previously mentioned, attributed the Higueruelas Fm to the Tithonian and the Villar del Arzobispo Fm to the Late TithonianMiddle Berriasian
, in an attempt to adapt the Boreal ages to Mediterranean stages, firstly and to the Global Time Scale, later (Fig.
Previous European reports include specimens from the Late Jurassic Solnhofen Limestone Formation (Tithonian) of southern Germany and equivalent-aged rocks of northern France (Bennett, 2007), the Purbeck Limestone Group (Berriasian
) of southern England (Howse and Milner, 1995), and the Camarillas Formation (lower Barremian) of Burgos, Spain (Sanchez-Hernandez et al., 2007) and the Leza Formation (Lower Cretaceous) of La Rioja, Spain (Pereda Suberbiola et al., 2012b).
"Thalassemys ruetimeyeri" Lydekker, 1889b, from the Berriasian
of Dorset, England; now recognized as a junior synonym of the pleurosternid Dorsetochelys typocardium (Seeley, 1869)) (Lapparent de Broin et al., 1996; Lapparent de Broin, 2001; Perez-Garcia, 2014; Perez-Garcia et al., 2015).
The dermal armour is very similar to that of goniopholidids such as Goniopholis simus from the Berriasian
of England and Germany, Anteophthalmosuchus hooleyi from the Valanginian of England, the unnamed Dollo's goniopholidid from the Barremian-Aptian of Belgium, Goniopholis baryglyphaeus from the Kimmeridgian of Portugal, and Sunosuchus junggarensis from the Late Jurassic of China (Wu et al., 1996; Salisbury and Frey, 2001; Schwarz, 2002; Salisbury and Naish, 2011).
Mantellisaurus atherfieldensis Barremian Delapparentia turolensis Hauterivian Valanginian Berriasian
Stage England Albian Aptian Iguanodon bernissartensis / Barremian Mantellisaurus atherfieldensis Barremian "Iguanodon seelyi" * Hauterivian Valanginian Hypselospinus fittoni Barilium dawsoni Kukufeldia tilgatensis * Sellacoxa pauli * Berriasian
Owenodon hoggii Stage References Albian McDonald et al.
g., Torcida Fernandez-Baldor et al., 2001, 2012; Castanera et al., 2010a; 2011) and the Berriasian