Taxones residentes Ordenes y familias de especies de insectos acuaticos encontrados: Coleoptera (larvas y adulto de; Curculionidae, Dytiscidae, Haliplidae, Hydrochidae, Hydrophilidae, Limnichidae, Luthrochidae y Noteridae), Ephemeroptera (larvas), Hemiptera (larvas y adultos de; Belostomatidae, Corixidae, Hebridae, Hydrometridae, Gerridae, Macroveliidae
, Mesoveliidae, Naucoridae, Nepidae, Notonectidae, Veliidae), Megaloptera (larvas de Corydalidae), Plecoptera (larvas), Trichoptera (larvas) Odonata (larvas).
Polhemus, 2008a): Mesoveloidea: Mesoveliidae: Madeoveliinae, Mesoveliinae (12 genera: 46 species), Hebroidea: Hebridae: Hebrinae, Hyrcaninae (7 genera: 126 species), Hydrometroidea: Hydrometridae: Heterocleptinae, Hydrometrinae, Limnobatodinae (7 genera: 126 species), Paraphrynoveliidae (1 genus: 2 species), Macroveliidae (3 genera: 3 species), Gerroidea: Hermatobatidae (1 genus: 10 species), Veliidae: Haloveliinae, Microveliinae, Ocelloveliinae, Perittopinae, Rhagoveliinae, Veliinae (61 genera: 962 species), and Gerridae: Charmatometrinae, Cylindrostethinae, Eotrechinae, Gerrinae, Halobatinae, Ptilomerinae, Rhagadotarsinae, Trepobatinae (67 genera: 751 species).
Besides providing minimum ages for clades, fossils may provide valuable information on the origin of adaptive traits, e.g., mate-guarding (Andersen and Poinar, 1992), but assignments of fossils to taxa of a quite generalized heteropteran appearance, such as Mesoveliidae and Macroveliidae, remain dubious (Damgaard, 2008a).
1, and expanded with his hypotheses for relationships within the Hydrometridae (Andersen, 1977), Macroveliidae (Andersen, 1978), Hebridae (Andersen, 1981) and Mesoveliidae (Andersen, 1999a), and with more recently described genera assigned to these families tentatively placed among their presumed closest relatives (Zettel, 1999; Andersen and D.
On the other hand, the Gerromorpha contain six well-supported and/or well-diagnosed clades: Mesoveliidae, Hebridae, Hermatobatidae, Hydrometridae, Macroveliidae + Paraphrynoveliidae and Veliidae + Gerridae, five of which were already recognized by Andersen (1982).
The Macroveliidae and Paraphrynoveliidae are the smallest of all currently recognized families, the former, consisting only of Macrovelia hornii Uhler, 1872; Chepuvelia usingeri China, 1963 and Oravelia pege Drake and Chapman, 1963, and Paraphrynovelia brinckii, 1957 and P.
No macropterous form is known for any of the two species of Paraphrynovelia, and Damgaard (2008b) could not find any convincing morphological characters to diagnose Paraphrynoveliidae, whereas Macroveliidae was diagnosed by the presence of paired, longitudinal ridges of the basal abdominal tergites and two characters also found in Hydrometridae: Eyes distinctly removed from the anterior margin of prothorax and the dorsally placed metathoracic spiracle.