The best position for Canrightia in Piperales
, sister to Saururaceae and Piperaceae, is two or three steps less parsimonious.
The eumagnoliids are an informal group composed of the Canellales, Laurales, Magnoliales, Piperales
and Chloranthales (sensu Soltis et al., 2000).
Utility of ontogenetic and conventional characters in determining phylogenetic relationships of Saururaceae and Piperaceae (Piperales
The angiosperms closest to the origin of monocots include Chloranthaceae and the Piperales
(Aristolochiaceae, Lactoridaceae, Piperaceae, and Saururaceae), all of which do have vessels.
Fossil perianthless Piperales
: a saururaceous inflorescence and flowers with in situ pollen from the Princeton Chert [British Columbia].
This is not surprising, given the characters that "supported" the original group: "The other line, here called the 'paleoherbs,' is characterized by anomocytic stomata, two perianth cycles, and trimery in both the perianth and the androecium (except for loss of one or both perianth cycles in Lactoris and Piperales
and secondary multiplication of parts in Nympheaceae)" (Donoghue & Doyle, 1989a: 28).
P-type plastids of different forms prevail in the other eumagnoliid orders; namely, Laurales, Magnoliales, and Piperales
(see Behnke, 1988).
and the Monocots: Altemative hypotheses for the origin of monocotyledonous flowers.
Although it has been suggested that the Chloranthaceae are related to the Piperales
(e.g., Cronquist, 1981), this relationship has not been supported in recent cladistic analyses.
and the Monocots: alternate hypotheses for the origin of monocotyledonous flowers.
However, it is not homologous with the perisperms of other monocotyledons (or dicotyledons such as some Piperales
and Nymphaeales: Dahlgren & Clifford, 1982), which are not derived from the nucellar epidermis.