10) shared with Helwingia (ambiguous for Phyllonoma), (5) nested within Solanales as sister to the Montiniaceae (Fig.
With the fossil, this monophyletic group collapses and so does the Solanales, but the Lamiales only lose the first two diverging taxa, the remaining 29 still form a clade.
However one of the alternatives relates to the Solanales, not as sister to the Solanaceae but as sister to the Montiniaceae (Fig.
Later, Lindley, having placed the Apocynaceae and Asclepiadaceae in the Gentianales, proceeded to place them apart (Lindley, 1845), the Apocynaceae being in the Gentianales and the Asclepiadaceae in the Solanales.
Problems in assigning the monotypic Retziaceae to the Loganiaceae (Leeuwenberg & Leenhouts, 1980), Scrophulariales (Dahlgren, 1983), or Solanales (Cronquist, 1988) may be due to its origin at an early stage when these three orders were not as distinct as they are today.
Note the isolated position of the Menyanthaceae, confirming its exclusion from the Gentianales and possibly the Solanales and Scrophulariales as well.