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The following article is from The Great Soviet Encyclopedia (1979). It might be outdated or ideologically biased.



(stool out), the formation of aboveground shoots from a node located at the base of the principal shoot in grasses and some other plants. A tillering node consists of a number of short, neighboring internodes, from whose buds lateral shoots are formed. Daughter lateral shoots develop from the axillary buds of the sheath, which either remain inside the sheath (in-travaginal shoots) or pierce the sheath and emerge to the outside (extravaginal shoots). If the shoots grow at an angle to the main shoot, a loose shrub forms; if they grow vertically, a compact shrub develops. As a result of the development of numerous shoots from the tillering node, perennial plants form turf. Long shoots distributed horizontally form rootstocks when they are underground (for example, wheatgrass) or offshoots when they are aboveground, (for example, meadow grass and reed).

In annual plants, tillering occurs early in their life (in winter plants, during autumn and spring; in spring plants, during spring) and ends with their actual shooting. In perennial plants, tillering is interrupted only during flowering and fruit-bearing and continues until the end of the vegetative period. During tillering not all shoots flower and yield a harvest; some die off, not attaining full development.

A distinction is made between total frutescence (the total number of shoots in the bush) and productive frutescence (the number of fruit-bearing shoots). The productivity of a plant depends on the species, variety, germination conditions, and, in perennial plants, the age and ability to furnish reserve organic substances. The longer the tillering period, the greater the shoot formation. Tillering also depends on agricultural methods: the time and rate of sowing (early sowing usually causes greater tillering; in dense plantings there is less), seed quality (the larger the seed, the greater the tillering), depth of planting, and application of fertilizers. Intensified tillering, within certain limits, increases the harvest. Late-forming shoots (aftergrowth) do not yield a harvest and weaken the development of productive stems. In pastures and hay fields, tillering is regulated by applying fertilizers (mainly nitrogenous ones), changing the water regimen, and rotating crops.


The Great Soviet Encyclopedia, 3rd Edition (1970-1979). © 2010 The Gale Group, Inc. All rights reserved.
References in periodicals archive ?
Nevertheless, it was expected that the ATAp was greater at higher levels of N, considering the effects of this nutrient on the tillering of forage grasses.
Micronutrient application substantially improved the plant height and grain number per spike, which were maximum when 'Shelter' was applied at tillering, booting, jointing and earing stage followed by the plots receiving shelter at tillering, booting and jointing stage (Table I).
Effects of nitrogen supply on tillering dynamics and regrowth of perennial ryegrass populations.
The tillering order, in general, followed the expected pattern (Neuteboom and Lantinga 1989).
This has been associated with increased productivity in both cases (Hunt and Easton, 1989; Hazard and Ghesquiere, 1997), but contrary to previous indications (Robson, 1967; Zarrough et al., 1984), decreased AL and tillering were not necessarily associated with long leaf length.
By observing positive effects of thiourea on plants, this study was carried out to evaluate the response of late sown wheat to foliar applied thiourea at tillering, jointing and booting stages and optimized the suitable concentration of thiourea for better growth of late sown wheat.
These results confirmed the assertion that the level of radiation becomes more limiting to tillering than the availability of nutrients under conditions of excessive shading.
From transplanting to maximum tillering stage it was similar to other varieties but afterwards length increased at much faster rate than others so it became vulnerable to lodging.
To evaluate the population pattern of tillering, three PVC rings 10 cm in diameter (0.0078 [m.sup.2]) were used, fixed on the soil in each experimental unit.
Similar trend was noted in WITA-4 and FARO 44 rice varieties where the more tillers were produced on the crop, the more they compensated for the number of spikelets per spike because a high tillering rice plant tends to "feed" more than the low tillering one.
Keywords: Date of sowing, wheat varieties, germination, tillering and grain yield