Pleural field gently vaulted, with five pleural ribs evenly bent posteriorly adaxially
. Medial part of the pleural ribs covered with densely spaced large granules and small to medium-sized tubercles gradually fading outwards and inwards.
6b); in Heterocoma and Proteopsis they are more concentrated on the abaxial face and adaxially
near the ribs (amphy-hypostomatic leaves) (Fig.
Fragment 7.34 cm long of a pinnate leaf with rachis 4.11 mm wide, slightly carenate with wide transverse protuberances (0.37 mm of width), in which opposite leaflets of round section, parallel margins and base slightly expanded are adaxially
inserted at an angle of 85[degrees]-90[degrees].
Costae are quite prominent, adaxially
sulcate, rather curved abaxially, moderately scaly, and densely pilose.
Petiole with 2-3 grooves adaxially
; sori continuous along pinnae margins.
1997), the glabellar S3 and S2 apodemes reduced to a low knob in the axial furrow (seen ventrally), with no furrows continuing adaxially
onto the glabella on the dorsal side, the thorax with fewer segments (nine) but the additional preanterior pair of pleurae on the pygidium being remnant from the juvenile thorax, and the inner two pairs of pleurae reduced but the additional pair developed on the ventral side of the pygidium.
Longitudinal glabellar furrows slightly inclined anteriorly adaxially
, strongly impressed posteriorly, gradually fading anteriorly, terminated at short distance from the preglabellar furrow.
Petiole 3-12 mm; leaf obovate to ovate, 3-5.5 cm, thick papery, adaxially
pubescent to glabrous, abaxially, and glabrous.
3m, n), each subtending an ovuliferous scale consistently bearing two adaxially
placed ovules, their micropyles directed toward the cone axis.
Leaves widely obovate to widely depressed obovate, widely elliptic to oblate, widely ovate-circular, very widely ovate to widely depressed ovate, length/width ratios 1.2-0.8:1; apices commonly retuse, sometimes rounded; crenate-dentate; leaves when yet unfolded adaxially
subglabrous, with minute glandular hairs throughout, pubescent and tomentose over teeth, and puberulent here and there on surface, pubescent from a few teeth apices, tufted on midvein and some secondary veins, when newly unfolded adaxially
subglabrate except at apex and base; abaxially pubescent (rays planar), golden-tomentellous, later glabrate, glabrescent, gray trichomes, with primary to fifth order veins raised; white striae (wax?) along veins.