Calyx with unequal lobes, elliptic or elliptic-ovate, sericeous, glabrous or not at the base, ciliate at margin, calcarate
lobe can reach twice the size of lateral lobes base, straight spur at the young bud, curving during maturity 0.5-0.7 cm long, densely pilose.
str.) as calcarate. The polarized endothecial thickening of Mutisioideae is shared with Carduoideae and Cichorioideae, although in some tribes other types of thickening may exist (Bremer, 1994).
This hypothesis suggests that the ancestor was a woody plant with alternate leaves, large and discoid capitula, actinomorphic deeply five-lobed corollas, caudate and calcarate anthers with sterile apical appendages, shortly bilobed styles glabrous or papillose with continuous stigmatic areas, and a pappus of scabrid bristles.