As a component of a coadapted
complex, theory evolves in the direction of further justifying government's often increasing regulatory mandate for securing food safety.
This is contrary to models of speciation that rely on a random disruption of well coadapted
traits that are continually buffered by stabilizing selective pressures within species (Carson 1982, 1985).
This Evolutionary Hypothesis proposes that different coadapted
gene systems would have evolved independently in each parental genome.
Hayward and Breese (1993) described the genetic structure of inbred cereal landraces as "stable mixture of coadapted
genotypes having different and complementary requirements for resources such as nutrients and light, and thus escaping competitive elimination".
Long-distance dispersal may take the individual to a habitat to which it is less adapted (Dias and Blondel 1996) and/or cause it to produce less fit offspring when reproducing with a philopatric individual, due to break down of coadapted
gene complexes (Shields 1982; Waser and Price 1994).
Genetic revolution and the origin of coadapted
genomes in parthenogenetic strains.
Intentional inbreeding may then be favored by the evolution of coadapted
gene complexes and/or adaptations to local environmental conditions (Mayr 1963, Moll et al.
polymorphism; Wade 1998) or may fix the population for a single coadapted
set of maternal and offspring characters.
Since oviposition and gall formation is probably under the control of coadapted
multilocus genetic systems, large shifts are unlikely.
When populations are adapted to different local conditions, genetic admixture can result in outbreeding depression or a reduction in population fitness caused by a breakdown of coadapted
gene complexes or by swamping local adaptation (Shields 1982; Templeton 1986).
It is therefore unlikely that outbreeding depression due to disruption of coadapted
gene complexes (Templeton 1986; Potts et al.
The assumption was that differently coadapted
populations would always result in hybrids with reduced fitness.