9A and B) and Left Pes 4 (LP4; Fig.9C and D) are the clearest imprints and are pentadactyl. The manus is poorly preserved and or often absent.
The pes is pentadactyl, with increasing digit length from digits I through IV, and IV being the longest.
Throughout this paper we use the Savage and Lips (1993) concepts of Celestus (no claw sheath) and Diploglossus (claw sheath present) in referring to valid
pentadactyl species.
A trait like the
pentadactyl limb of tetrapods, however, seems to have an enormous amount of inertia.
Footprints of a quadrupedal tetrapod in which the manus is tetradactyl and the pes is
pentadactyl. The manus and pes are of approximately equal size (especially in widths), generally plantigrade and may be overstepped.
The track's features are typical of "lacertoid" footprints: a
pentadactyl manual print with digits increasing in length from I to IV.
He overlooks the pair PENTADACTYLE-PRECIPITANTS with a shift of 15, listed in my book Wordplay, A Curious Dictionary of Language Oddities (Sterling, 1999)." The first word is found in Webster's Second as a variant of
PENTADACTYL, having five fingers or toes.
DIAGNOSIS: Quadrupedal, plantigrade tetrapod trackway with tetradactyl manus and
pentadactyl pes imprints.
This material includes well preserved
pentadactyl and tetradactyl trackways of at least six different forms of both small and rather large tetrapods that were capable of locomotion on land (Smithson et al.
Description: Marius and pes are of similar proportions, both
pentadactyl, with digits increasing in length from I to IV, and digit V somewhat offset.