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The transition between the basal part of the perforatorium and the anterior part of the nucleus was visible (approximately 25 nm).
En esta etapa se extruye en forma explosiva el perforatorium que se encuentra en el interior del tubo acrosomico formando una estructura filiforme en medio de la abertura anterior del acrosoma, el filamento acrosomico.
Kristensen (1991) listed for this clade the following putative synapomorphies: maxillary lacinia elongate and slender, detached from stipes; postclypeus enlarged; reduced number of tarsomeres (three or less); cerci absent; at most six Malpighian tubules; abdominal ganglia fused in one mass; gonangulum in the females fused with tergum nine (Scudder 1961); and sperm acrosome without perforatorium (Jamieson et al.
The acrosomal complex can be divided into an acrosomal vacuole, amorphous subacrosomal material and the acrosomal filament (perforatorium) starting from the amorphous subacrosomal material.
Firstly, there is no extrusion of an acrosomal process or "perforatorium" by the polymerization of actin, as occurs throughout molluscs and other metazoan groups (see review by Tilney, 1985).
Neither a distinct perforatorium rod nor filamentous structures are detected within the subacrosomal material.
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