The dorsal surface of the skull is covered with dermal tuberosities but the jugal, the quadratojugal, the maxilla, the premaxilla and the occipital parts of squamosal are free from sculpturing (Figs 1, 2).
The lateral temporal region is slightly damaged in PIN 4659/1 but there appears to be an emargination between the jugal and the quadratojugal (Fig.
Maxilla and quadratojugal separated or in contact (22)
Tsuji (2006) scored the maxilla and the quadratojugal as being in contact in Nyctiphruretus based on illustrations of Ivakhnenko (1979) and Lee (1995, 1997), when in fact they are not due to a temporal emargination between the quadratojugal and the jugal (Fig.
Quadratojugal lateral temporal fenestra contribution (32)
Squamosal delicate; anterior arm slender; posterior arm three fifths length of anterior arm, medial ala overlying dorsolateral surface of crista parotica; ventral arm well developed, distally articulating with quadratojugal
laterally and posterior ramus of pterygoid medially.